They represent the most ancient and complex system of relationships between nature, historical, social, and cultural phenomena. The presence of human settlements and production activities as a continuous presidium of the territory, as a balanced use of the resources, and as a cultural richness is one of the essential features of the activities in the nature reserve. The towns, villages, bridges, mills, hermitages, fortresses, farmsteads, mule tracks, and carriage roads are signs of the territory witnessing the big or small events that have characterized the life of the local people. They represent a great local heritage which, if rediscovered, can be transformed into a precious resource to enhance the social and cultural image of the towns of the nature reserve.